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character: Tortoises
topic: Habitat and Role in Ecosystem
subject: Geography
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character: Desert Tortoise
topic: Habitat Adaptations and Ecosystem Impact
subject: Geography
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character: Galápagos Tortoise
topic: Influence on Island Ecosystem Dynamics
subject: Geography
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character: Pollinators
topic: Interactions with Tortoises in Their Habitat
subject: Geography
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